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Storm Reid - Meg, Oprah Winfrey - Mrs. Which, Reese Witherspoon - Mrs. Whatsit, Mindy Kaling - Mrs. Who, Levi Miller - Calvin, Deric McCabe - Charles Wallace, Chris Pine - Mr. Murry, Gugu Mbatha-Raw - Mrs. Murry, Zach Galifianakis - Happy Medium, Michael Peña - Red, André Holland - Principal Jenkins, Rowan Blanchard - Veronica Kiley, Bellamy Young - Camazotz Woman, David Oyelowo - The It (voice) , Conrad Roberts - Elegant Man