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Tom Holland - Ian Lightfoot (voice) , Chris Pratt - Barley Lightfoot (voice) , Julia Louis-Dreyfus - Laurel Lightfoot (voice) , Octavia Spencer - The Manticore (voice) , Mel Rodriguez - Colt Bronco (voice) , Kyle Bornheimer - Wilden Lightfoot (Dad) (voice) , Lena Waithe - Officer Spector (voice) , Ali Wong - Officer Gore (voice) , Tracey Ullman - Grecklin (Pawn Shop Owner) (voice) , Wilmer Valderrama - Gaxton (College Friend) (voice) , George Psarras - Officer Avel / Additional Voices (voice) , John Ratzenberger - Construction Worker Fenwick (voice)