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Jason Alexander - Father (voice) , Mary Steenburgen - Mother (voice) , Reese Witherspoon - Serena (voice) , Seth Green - Boyd (voice) , Carol Burnett - Mrs. Hammerbotham (voice) , Joe Mantegna - Monty (voice) , Sam Gifaldi - Sam Beaver (voice) , Dee Bradley Baker - Louie (voice) (as Dee Baker) , Melissa Disney - Billie (voice) , Kath Soucie - Serena (cygnet) / Paramedic / Newscaster (voice) , Elizabeth Daily - Ella (voice) (as E.G. Daily) , Pamela Adlon - A.G. Skinner (voice) (as Pamela Segall Adlon) , Steve Vinovich - Maurice / Ranger (voice) , Gary Anthony Williams - Sweets (voice) , Corey Burton - Senator (voice)